PD Dr. Alexander Kupfer


Zeitschriftenbeiträge (peer-reviewed, ISI-evaluated, SCOPUS-covered)
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Pogoda P, Kupfer A (2019). High osteological variation in a terrestrial salamander (genus Salamandrina). Zoologischer Anzeiger.

von Bülow B & Kupfer A (2019). Monitoring population dynamics and survival of Northern Crested Newts (Triturus cristatus) for 19 years at a pond in Central Europe. Salamandra 55: 97-102.

Geissler, P, Kupfer A (2019). On the taxonomic status of Lygosoma (Keneuxia) dubium Werner, 1909. Zootaxa 4543: 295–300.

Pogoda P, Kupfer A (2018). Flesh and bone: An integrative approach towards sexual size dimorphism of a terrestrial salamander (genus Salamandrina). Journal of Morphology 279: 1468–1479. abstract

Jared C, Mailho-Fontana PL, Jared SGS, Kupfer A, Delabie JHC, Wilkinson M, Antoniazzi MM (2018). Life history and reproduction of the neotropical caecilian Siphonops annulatus (Amphibia, Gymnophiona, Siphonopidae), with special emphasis on parental care. Acta Zoologica. abstract

Wartenberg N, Reinhard S, Nöllert A, Staniczek AH, Kupfer A (2017). Caddisfly larvae (Trichoptera: Phryganeidae) as scavengers of carcasses of the common frog Rana temporaria (Amphibia: Ranidae). Salamandra 53: 458–460. open access

Irisarri I, Baurain D, Brinkmann H, Delsuc F, Sire J-Y, Kupfer A, Petersen J, Jarek M, Meyer A, Vences M, Philippe H. (2017). Phylotranscriptomic consolidation of the jawed vertebrate timetree. Nature Ecology & Evolution 1: 1370-1378. abstract 

Kupfer A, Maxwell E, Reinhard S & S Kuehnel (2016). The evolution of parental investment in caecilian amphibians: a comparative approach. Biological Journal of the Linnean Society 119:4-14. abstract

Maerker M, Reinhard S, Pogoda P & A Kupfer (2016). Sexual size dimorphism in the viviparous caecilian amphibian Geotrypetes seraphini seraphini (Gymnophiona: Dermophiidae) including an updated overview of sexual dimorphism in caecilian amphibians. Amphibia-Reptilia 37:291-299. abstract

Budzik KA, Zuwala K, Kupfer A, Gower DJ & M Wilkinson (2015). Diverse anatomy of the tongue and taste organs in five species of caecilian (Amphibia: Gymnophiona). Zoologischer Anzeiger 257:103-109. abstract

Schoch R, Poschmann M & A Kupfer (2015). The salamandrid Chelotriton paradoxus from Enspel and Randeck Maar (Oligocene-Miocene), Germany. Palaeobiodiversity and Palaeoenvironments 95: 77-86. abstract 

Reinhard S & A Kupfer (2015). Sexual dimorphism in a French population of the Marbled Newt Triturus marmoratus (Urodela: Salamandridae). Salamandra 51: 121-128. open access

Geissler P, Poyarkov Jr. NA, Grismer L, Nguyen TQ, An HAT, Neang T, Kupfer A, Ziegler T, Böhme W & H Müller (2015). New Ichthyophis species from Indochina (Gymnophiona, Ichthyophiidae): 1. The unstriped forms with descriptions of three new species and the redescriptions of I. acuminatus Taylor, 1960, I. youngorum Taylor, 1960 and I. laosensis Taylor, 1969. Organsims, Diversity & Evolution 15: 143-174. abstract

Reinhard S, Renner S & A Kupfer (2015). Age and sexual dimorphism in Mediterranean salamanders. Zoology 118: 19-26. abstract

Reinhard S, Renner S & A Kupfer (2015). Age and fecundity in Salamandra algira. Salamandra 51:19–24.

Reinhard S, Voitel S, Kupfer A (2013). Paternal care in a neotenic salamander. Zoologischer Anzeiger 253: 1-5. abstract

Kuehnel S & A Kupfer (2012). Sperm storage in caecilian amphibians. Frontiers in Zoology. 9. open access

Kuehnel S, Herzen J, Kleinteich T, Beckmann F & A Kupfer. (2012). Functional reproductive morphology: the female cloaca of an oviparous caecilian amphibian. Acta Zoologica 93: 208-221. abstract

Scholz S, Orlik M, Gonwuo LN & A Kupfer (2010). Demography and life history of a viviparous Central-African caecilian amphibian. Journal of Zoology 280: 17-24. abstract

Kupfer A (2009). Sexual size dimorphism in caecilian amphibians: analysis, review and future directions. Zoology 112: 362-369. abstract

Kupfer A, Kühnel S, Vetter J, Olsson L. (2009). Reproductive and developmental biology of caecilian amphibians. Integrative and Comparative Biology 49(Suppl 1): e95. 

Kupfer A, Wilkinson M, Gower DJ, Müller H & R Jehle (2008). Care and parentage in a skin-feeding caecilian amphibian. Journal of Experimental Zoology 309A: 460-467. abstract

Wilkinson M, Kupfer A, Marques-Porto R, Jeffkins H,  Antoniazzi MM, Jared C (2008). One hundred million years of skin feeding? extended parental care in a Neotropical caecilian (Amphibia: Gymnophiona). Biology Letters 4: 358-361. abstract

Kupfer A, Müller H, Antoniazzi MM, Jared, C, Greven H, Nussbaum RA & M Wilkinson (2006). Parental investment by skin feeding in a caecilian amphibian. Nature 440: 926-929. abstract   watch skin feeding on you tube

Kupfer A, Langel R, Scheu S, Himstedt W & M Maraun (2006). Trophic ecology of a tropical aquatic and terrestrial food web: insights from stable isotopes (15N). Journal of Tropical Ecology 22:469-476. abstract

Kupfer A, Gaucher P, Wilkinson M & DJ Gower (2006). Passive trapping of aquatic caecilians (Amphibia: Gymnophiona: Typhlonectidae). Studies in Neotropical Fauna and Environment 41: 93-96. abstract

Kupfer A, Kramer A, Himstedt W & H Greven (2006). Copulation and egg retention in a terrestrial caecilian (genus Ichthyophis). Zoologischer Anzeiger 244: 223-228. abstract

Kupfer A, Nabithabatha J & W Himstedt (2005). Life history of amphibians in the seasonal tropics: habitat parameters, community ecology and population structure of a caecilian (genus Ichthyophis). Journal of Zoology 266: 237-247. abstract

Kupfer A, Nabithabatha  J & W Himstedt (2005). From water into the soil: trophic ecology of a caecilian amphibian (genus Ichthyophis). Acta Oecologica 21: 95-105. abstract

Montero I, Reichle S & A Kupfer (2005). Observations on the reproductive ecology of Siphonops paulensis Boettger, 1892 (Gymnophiona: Caeciliidae) in Bolivia. Salamandra 41: 91-94. open access

Kupfer A, Nabithabatha J & W Himstedt (2004). Reproductive ecology of female caecilian amphibians (genus Ichthyophis): a baseline study. Biological Journal of the Linnean Society 83: 207-217. abstract

Kupfer A & H Müller (2004). On the taxonomy of ichthyophiid caecilians from southern Thailand: A reevaluation of the holotype of Ichthyophis supachaii Taylor 1960 (Amphibia: Gymnophiona: Ichthyophiidae). Amphibia-Reptilia 25: 87-97. abstract

Kupfer A, Kramer A & W Himstedt (2004). Sex related growth patterns in a caecilian amphibian (genus Ichthyophis). Journal of Zoology 262: 173-178. abstract

Kupfer A, Gower DJ & W Himstedt (2003). Field observations on the predation of the caecilian amphibian, genus Ichthyophis (Fitzinger, 1926) by the red-tailed pipe snake Cylindrophis ruffus (Laurenti, 1768). Amphibia-Reptilia 24: 212-215. abstract

Gower DJ, Kupfer A, Oommen VO, Himstedt W, Nussbaum RA, Loader SP, Presswell B, Krishna SB, Müller H, Boistel R & M Wilkinson (2002). A molecular phylogeny of ichthyophiid caecilians (Amphibia: Gymnophiona: Ichthyophiidae): Out of India or out of South East Asia? Proceedings of The Royal Society London B Biology 269: 1563-1569. abstract

Kupfer A & SFM Teunis (2001). Field data on antipredator responses in three species of newts (genus: Triturus). Herpetological Journal 11: 117-120.

Kupfer A & S Kneitz (2000). Population ecology of Great Crested Newts (Triturus cristatus) in an agricultural landscape: dynamics, pond fidelity and dispersal. Herpetological Journal 10: 165-172.


Zeitschriftenbeiträge (peer-reviewed, non ISI-evaluated, Auswahl)

Penell A & A Kupfer (2018). Phänotypische Bestimmung von Kleinen Wasserfröschen und Teichfröschen (Pelophylax lessonae und P. „esculentus“) aus dem westlichen Federsee in Südwestdeutschland. Rana 19: 74-85.

Kupfer A & U Hoßfeld (2013). Fortpflanzungstrategien der Amphibien. Praxis der Naturwissenschaften/ Biologie in der Schule 63: 36-40.

Kozlik M, Reinhard S, Kupfer A & Hoßfeld (2013). Warum Kriechtiere heute Sauropsida heißen sollten! MNU 66: 113-115.

Haacks M, Bock D, Drews A, Flottmann H-J, Geske C, Kupfer A, Ortmann D & R Podloucky (2009). Bundesweite Bestandserfassung von Kammmolchen im Rahmen des FFH-Monitorings. Erfahrungen zur Fängigkeit von verschiedenen Wasserfallentypen. Natur & Landschaft 84: 276-280.

Kupfer A & M Maraun (2003). Natural History Notes: Ichthyophis kohtaoensis (Koh-Tao Caecilian). Diet. Herpetological Review. 34: 226.

Kupfer A (1999). Eine schnelle Methode zur Längenmessung von Kammolchen (Triturus cristatus) im Freiland. Zeitschrift für Feldherpetologie 6: 233-234.

Kupfer A (1998). Wanderstrecken einzelner Kammolche (Triturus cristatus) in einem Agrarlebensraum. Zeitschrift für Feldherpetologie 5: 238-242. 

Breuckmann A & A Kupfer (1998). Zur Umsiedlung einer Kammolchpopulation (Triturus cristatus) im nordöstlichen Ruhrgebiet: ein Rückblick nach zehn Jahren. Zeitschrift für Feldherpetologie 5: 209-218. 

Mölle J & A Kupfer (1998). Amphibienfang mit der Auftauchfalle: Methodik und Evaluierung im Freiland. Zeitschrift für Feldherpetologie 5: 219-228. 

Kupfer A (1997). Phänologie und Metamorphosegrößen juveniler Kammolche, Triturus cristatus: ein Vergleich von zwei benachbarten Populationen. Zeitschrift für Feldherpetologie 4: 141-155. 

Vences M, Kupfer A, Llorente GA, Montori A & MA Carretero (1997). Description of the larval stages of the Pyrenean frog, Rana pyrenaica Serra-Cobo, 1993 (Amphibia: Ranidae). Bolletino del Museo Regionale di Scienze Naturali Torino 15: 1-23.

Kupfer A (1995). Bemerkungen zum Abwehrverhalten des Bergmolches Triturus alpestris alpestris im Freiland. Zeitschrift für Feldherpetologie 2: 85-90. 



Thiesmeier B, Kupfer & Jehle R (2009): Der Kammmolch - Ein Wasserdrache in Gefahr, 2nd. Edition. Bielefeld (Laurenti).

Thiesmeier B & A Kupfer (2000): Der Kammmolch - Ein Wasserdrache in Gefahr. Bochum (Laurenti).



Kupfer A B v Bülow (2011). Nördlicher Kammmolch. In Arbeitskreis Amphibien Reptilien NRW (Hrsg.). Handbuch der Amphibien und Reptilien Nordrhein-Westfalens, pp. 376-406. Laurenti, Bielefeld.

Kühnel S, Reinhard S & A Kupfer (2010). Evolutionary reproductive morphology of amphibians: an overview. Bonn Zoological Bulletin 57: 119-126.

Schlüpmann M & Kupfer A (2009). Methoden der Amphibienerfassung: ein Überblick. In Hachtel M, Schlüpmann M, Thiesmeier B & K Weddeling (Hrsg.). Methoden der Feldherpetologie. Supplement der Zeitschrift für Feldherpetologie 15: 7-84. 

Kupfer A (2007). Sexual size dimorphism in amphibians: an overview. In Fairbairn DJ, Blanckenhorn WU & T Szekely (eds) Sex, size and gender roles: evolutionary studies of sexual size dimorphism. Oxford, Oxford University Press. abstract

Kupfer A (2006). Caecilians. In Brockhaus Animal Enzyclopedia. Volume 4, Amphibians & Reptiles, pp. 158-171. Bibliographisches Institut & F.A. Brockhaus AG, Mannheim (in German). 

Kupfer A (2001). A review on the monitoring methods for great crested newts (Triturus cristatus). In Krone A (ed) The great crested newt (Triturus cristatus): ecology and distribution. Natur und Text, Rangsdorf (in German with English summary).

Kupfer A & B v Bülow (2001). Great crested newts (Triturus cristatus) in Nordrhine-Westfalia: distribution, habitats und threats. In Krone A (ed) The great crested newt (Triturus cristatus): ecology and distribution, pp. 83-91. Natur und Text, Rangsdorf (in German with English summary).

Kupfer A & S Kneitz (1999). Colonisation processes and population dynamics of amphibians in an agricultural landscape. In Boothby J (ed) Ponds and pond landscapes of Europe, pp. 161-168. John Moores University, Liverpool