Dr. Arnaud Faille

Profilbild Arnaud Faille

Curator & Deputy head of Department Entomology



+49 (0) 711 / 8936-221
+49 (0) 711 / 8936-100

Research interests

Evolutionary biology, phylogenetics, systematic and phylogeny of Coleoptera, origin and evolution of biodiversity, evolution of endogean and cave beetles, phylogeny vs morphological evolution in beetles, biogeography of beetles, adaptive and speciation processes, conservation of subterranean biodiversity.

Target taxa: Cave Invertebrates, Insects (Coleoptera).




  • 81. Faille A. 2021. Chapter 6: Cave biogeography. 143-164. In: Guilbert E. (Ed.): Biogeography: an integrative approach of the evolution of living. CNRS. ISTE Ltd/ John Wiley & Sons, Inc : London, Hoboken.

  • 80. Faille A., Deharveng L. 2021. The Coume Ouarnède System, a Hotspot of Subterranean Biodiversity in Pyrenees (France). Diversity 2021, 13, 419. https://doi.org/10.3390/d13090419

  • 79. Wynne JJ, Howarth FG, Mammola S, Ferreira RL, Cardoso P, Di Lorenzo T, Galassi DMP, Medellin RA, Miller BW, Sánchez-Fernández D, Bichuette ME, Biswas J, Blackeagle CW, Boonyanusith C, Borges PAV, Boston PJ, Cal RN, Cheeptham N, Deharveng L, Eme D, Faille A, Fenolio D, Fišer C, Fišer Ž, ʻOhukaniʻōhiʻa Gon III SM, Goudarzi F, Griebler C, Halse S, Hoch H, Kale E, Katz AD, Kováč L, Lilley TM, Manchi S, Manenti R, Martínez A, Meierhofer MB, Miller AZ, Moldovan OT, Niemiller ML, Peck SB, Pipan T, Pellegrini TG, Phillips-Lander CM, Poot C, Racey PA, Amorim de Rosário I, Sendra A, Shear WA, Souza Silva M, Taiti4 S, Tian M, Venarsky MP, Pakarati SY, Zagmajster M, Zhao Y. A Roadmap for the Conservation of the Subterranean Biome. Conservation Letters. 2021;e12834.

  • 78. Kavanaugh DH, Maddison DR, Simison WB, Schoville SD, Schmidt J, Faille A, Moore W, Pflug JM, Archambeault SL, Hoang T, Chen JY. 2021. Phylogeny of the supertribe Nebriitae (Coleoptera: Carabidae) based on analyses of DNA sequence data. Zookeys. 1044: 41–152.

  • 77. Huang S., Zhou J., Tian M. & Faille A. 2021. Three new species of the subterranean genus Guiodytes from Guangxi, China (Coleoptera: Carabidae: Clivinini). European Journal of Taxonomy 774: 135–154. https://doi.org/10.5852/ejt.2021.774.1537

  • 76. Faille A., Balart-García P., Fresneda J., Bourdeau C., Ribera I. 2021. A remarkable new genus of Iberian troglobitic Trechodina (Coleoptera: Carabidae: Trechinae: Trechini), with a revisited molecular phylogeny of the subtribe. Annales de la Société Entomologique de France. 57(2): 85-106.


  • 75. Huang, S., Tian, M., Faille, A. 2020. Two new species of subterranean ground beetles from China (Coleoptera: Carabidae: Trechinae). Raffles Bulletin of Zoology. Suppl. 35. 22-31. (See also: Raffles Bulletin of Zoology, 68: 237 (23.IV.2020), doi: 10.26107/RBZ-2020-0020: Erratum to Huang SB, Tian MY & Faille A (2020) Two new species of subterranean ground beetles from China (Coleoptera: Carabidae: Trechinae). Raffles Bulletin of Zoology, Supplement 35: 22–31.)
  • 74. Huang, S., Tian, M., Faille, A. 2020. Three new species of the aphaenopsian trechine genus Pilosaphaenops Deuve & Tian, 2008 from South China Karst (Coleoptera: Carabidae: Trechinae).Annales de la Société Entomologique de France (n.s.). 56:3, 203-214, doi: 10.1080/00379271.2020.1771203
  • 73. Huang S., Tian M., Faille A. 2020. A preliminary phylogeny of cave trechine beetles from China (Coleoptera: Carabidae: Trechini). Proceedings of the 25th International Conference on Subterranean Biology (Alba Iulia, 28th June – 4th July 2020). ARPHA Conference Abstracts 3: e518973: e51897 doi: 10.3897/aca.3.e51897
  • 72. Sendra A., Antić D., Barranco P., Borko Š., Christian E., Delić T., Fadrique F., Faille A., Galli L., Gasparo F., Georgiev D., Giachino P.M., Kováč L., Lukić M., Marcia P., Miculinić K., Nicolosi G., Palero F., Paragamian K., Pérez T., Polak S., Prieto C.E., Turbanov I., Vailati D. & Reboleira A.S.P.S. 2020. Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae). European Journal of Taxonomy 591: 1–138.


  • 71. Moret P, Barragán A, Murienne J, Faille A, Gobbi M. 2019. High altitude endemic carabid beetles as biogeographic indicators: a case study in equatorial Andes. Carabids in extreme environments. 19th European Carabidologists Meeting. ARPHA Conference Abstracts 2: e38693. https://doi.org/10.3897/aca.2.e38693
  • 70. Huang, S., Tian, M., Faille, A. 2019. The most remarkable cave-specialized trechine beetles from China. Carabids in extreme environments. 19th European Carabidologists Meeting. ARPHA Conference Abstracts 2: e37360 doi: 10.3897/aca.2.e37360
  • 69. Kavanaugh D, Maddison D, Simison B, Schoville S, Schmidt J, Faille A. 2019. Phylogeny of the supertribe Nebriitae (Coleoptera: Carabidae) based on analyses of molecular sequence data - some surprises. Carabids in extreme environments. 19th European Carabidologists Meeting. ARPHA Conference Abstracts 2: e37156 doi: 10.3897/aca.2.e37156
  • 68. Huang, S., Tú, V.T., Phú, P.V., Tian, M., Faille, A. 2019. Contributions to the knowledge of the genus Tonkinaphaenops Deuve, 2013 from China and Vietnam (Coleoptera: Carabidae: Trechinae). Zootaxa. 4701 (1): 35–53.
  • 67. Maghradze, E., Faille, A., Barjadze, S., Hlaváč, P. 2019. A new cavernicolous species of the genus Bergrothia Reitter, 1884 (Coleoptera, Staphylinidae, Pselaphinae) from Georgia. Zootaxa. 4608 (2): 371–379.
  • 66. Faille, A. 2019. Chapter 14 : Beetles. In : Encyclopedia of caves. Third Edition. White WB, Culver DC, Pipan T. (Eds.). Elsevier Inc. 102-108.
  • 65. Fresneda, J., Valenzuela, E., Bourdeau, C., Faille, A. 2019. Nouvelles espèces de Trechus Clairville, 1806 du clade de T. saxicola des Monts cantabriques, Asturies, Espagne. Biogéographie et spéciation. (Coleoptera, Carabidae, Trechinae). Annales de la Société Entomologique de France (n.s.). doi.org/10.1080/00379271.2018.1546553.
  • 64. Fresneda J., Faille A., Fery H., Ribera I. 2019. A molecular phylogeny of Speonemadus Jeannel 1922, with description of two new species from Morocco (Coleoptera : Leiodidae : Cholevinae : Anemadini). Zootaxa. 4543 (1): 1–36.
  • 63. Maddison D. R., Kanda K., Boyd O. F., Porch N., Faille A., Erwin T. L., Roig-Juñent S. 2019. Phylogeny of the beetle supertribe Trechitae (Coleoptera: Carabidae): unexpected clades, isolated lineages, and morphological convergence. Molecular Phylogenetics and Evolution. 132: 151-176.


  • 62. Hlaváč, P., Faille, A. 2018. New species of cavernicolous Tychobythinus Ganglbauer, 1896 (Coleoptera, Staphylinidae, Pselaphinae) from Turkey. Entomologische Blätter und Coleoptera. Papers celebrating the 80th birthday of Dr. Ivan Löbl. 114 : 233-236.
  • 61. Faille, A., Casale, A., Hernando, C., Aït Mouloud, S., Ribera, I. 2018. Tectonic vicariance versus Messinian dispersal in western Mediterranean ground beetles. Zoologica Scripta. 47 :565-581.
  • 60. Schmidt, J., Faille, A. 2018. Revision of Trechus Clairville, 1806 of the Bale Mountains and adjacent volcanos, Ethiopia (Coleoptera, Carabidae, Trechini). European Journal of Taxonomy. 446 : 1-82. doi.org/10.5852/ejt.2018.446
  • 59. Ribera, I, Cieslak, A, Faille, A, Fresneda, J. 2018. Chapter 10 : Historical and ecological factors determining cave diversity. in: Moldovan O, Kovác L, Halse S.(Eds) Cave Ecology. Ecological Studies, Springer. 229-252.
  • 58. Chłond, D., Guilbert, E., Faille, A., Baňař P., Davranoglou L.R. 2018. An amazing cave species of Madagascan Emesinae (Hemiptera: Heteroptera: Reduviidae). Zootaxa. 4425(2): 372-384.
  • 57. Thomson SA, Pyle RL, Ahyong ST, Alonso-Zarazaga M, Ammirati J, Araya JF, et al. (2018) Taxonomy based on science is necessary for global conservation. PLoS Biology 16(3): e2005075. doi.org/10.1371/journal.pbio.2005075
  • 56. Faille, A., Lecoq, J.C. 2018. Oedichirus spelaeus n.sp., the first cave dwelling beetle of Madagascar (Coleoptera : Staphylinidae : Paederinae). Journal of Cave and Karst studies. 80(1) : 13-18.
  • 55. Sánchez-Fernández D., Rizzo, V., Bourdeau, C., Cieslak, A., Comas, J., Faille, A., Fresneda, J., Lleopard, E., Millán, A., Pallarés, S., Pastor, J., Ribera, I. 2018. The deep subterranean environment as a model system in ecological, biogeographical and evolutionary research. Subterranean Biology. 25 : 1-7.


  • 54. Löbl, I., Faille, A. 2017. Toxidium cavicola sp. nov., a new cave dwelling Malagasy Scaphidiinae (Coleoptera: Staphylinidae). Annales Zoologici. 67(2) : 345-348.
  • 53. Hoch, H., Bourgoin, T., Faille, A., Soulier-Perkins, A. 2017. A new species of Bennini (Hemiptera, Cixiidae) from granitic subterranean environment in Vietnam, an under-investigated biotope. Annales Zoologici. 67(1): 1-12.
  • 52. Ortuño, V.M., Sendra, A., Reboleira, A.S.P.S., Fadrique, F., Faille, A. 2017. The Iberian genus Paraphaenops Jeannel, 1916 (Coleoptera: Carabidae: Trechini). Zoologischer Anzeiger - A Journal of Comparative Zoology. 266: 71-88.


  • 51. Faille, A., Ribera, I., Fresneda J. 2016. On the genus Aphaobius Abeille de Perrin, 1878, with description of a new species from the mesovoid shallow substratum (MSS) of Austria (Coleoptera: Leiodidae: Cholevinae: Leptodirini). Zootaxa. 4169(1): 44-56.
  • 50. Sánchez-Fernández, D., Rizzo, V., Cieslak, A., Faille, A., Fresneda, J., Ribera, I. 2016. Thermal niche estimators and the capability of poor dispersal species to cope with climate change. Scientific Reports. 6:23381 | DOI: 10.1038/srep23381
  • 49. Andújar, C., Faille, A., Pérez, S., Zaballos, J., Vogler, A., Ribera, I. 2016. Gondwanian relicts and oceanic dispersal in a cosmopolitan radiation of euedaphic ground beetles. Molecular Phylogenetics and Evolution. 99: 235-246.


  • 48. Faille, A., Bourdeau, C., Bellés, X., Fresneda, J. 2015. Allopatric speciation illustrated: The hypogean genus Geotrechus Jeannel, 1919 (Coleoptera: Carabidae: Trechini), with description of four new species from the Eastern Pyrenees (Spain). Arthropod Systematics & Phylogeny. 73(3): 439–455.
  • 47. Schmidt, J., Faille, A. 2015. Confirmation of occurrence of Trechus Clairville, 1806 in the Baltic amber forests, with description of a flightless edaphic species, and remarks on Trechoides Motschulsky, 1856 (Coleoptera: Carabidae: Trechini). Zootaxa. 4028(4): 539–550.
  • 46. Faille, A., Tänzler, R., Toussaint, E.F.A. 2015. On the way to speciation: Shedding light on the karstic phylogeography of the micro-endemic cave beetle Aphaenops cerberus in the Pyrenees. Journal of Heredity. 106(6): 692–699.
  • 45. Perreau, M., Faille, A. 2015. A new Ptomaphaminus Perreau, 2000 from a granitic subterranean environment in Vietnam (Coleoptera: Leiodidae: Cholevinae: Ptomaphagini). Zootaxa. 4021(1): 195–200.
  • 44. Fresneda, J., Alfambra, F., Bourdeau, C., Faille, A. & Ribera, I. 2015. Nuevas capturas de Aphaenops mensioni Lagar, 1976, con un inventario de la fauna hipogea terrestre del Parque Nacional de Ordesa y Monte Perdido. Exploracions. 21: 61–68.
  • 43. Fresneda, J., Valenzuela, E., Faille, A. 2015. Un nuevo Trechus Clairville, 1806 hipogeo troglomorfo de Asturias, España (Coleoptera, Carabidae, Trechinae). Bulletin de la Société entomologique de France. 120(3): 403–409.
  • 42. Faille, A., Pluot-Sigwalt, D. 2015. Convergent evolution in the reduction of ovariole number associated with subterranean life in cave beetles. PLOS ONE 10(7): e0131986. doi:10.1371/journal.pone.0131986.
  • 41. Fresneda, J., Bourdeau, C., Faille, A., 2015. Una nueva especie troglobiomorfa de Trechus Clairville, 1806 y evidencias de colonizaciones múltiples del medio subterráneo de los Montes Cantábricos (Coleoptera, Carabidae, Trechinae). Animal Biodiversity and Conservation, 38(1): 87–100.
  • 40. Faille, A. 2015. Biospeology: diversity, distribution and interest of cave biota. Proceedings of the 1st International Convention of Speleology. Barcelone, Spain, 1-3.V.2015. Pp. 25–32.
  • 39. Faille, A., Bourdeau, C., Deharveng, L. 2015. Weak impact of tourism activities on biodiversity in a subterranean hotspot of endemism and its implications for the conservation of cave fauna. Insect Conservation and Diversity. 8: 205–215.


  • 38. Faille, A., Comas, J. 2014. Nuevos datos y descripción del macho de Aphaenops mensioni Lagar 1976, una especie endémica del Parque Nacional de Ordesa y Monte Perdido (Coleoptera: Carabidae: Trechinae). Heteropterus. 14(2): 119–123.
  • 37. Faille, A., Andújar, C., Fadrique, F., Ribera, I. 2014. Late Miocene origin of a Ibero-Maghrebian clade of ground beetles with multiple colonisations of the subterranean environment. Journal of Biogeography. 41: 1979–1990.
  • 36. Deuve T., Faille, A. 2014. Structure secondaire de l’ARN 18S et phylogénie basale du genre Carabus Linné, 1758 (Coleoptera, Carabidae). Annales de la Société Entomologique de France (N.S.). 49(4): 430–445.


  • 35. Faille, A., Casale, A., Balke, M., Ribera, I. 2013. A molecular Phylogeny of Alpine subterranean Trechini (Coleoptera: Carabidae). BMC Evolutionary Biology. 13: 248.
  • 34. Faille, A., Fresneda, J., Bourdeau, C. 2013. Discovery of a new blind ground beetle in western french Pyrenees, and its relevance to the phylogeny of Pyrenean hypogean Trechini. Zootaxa. 3686(4): 482–492.


  • 33. Tian M., Faille, A. 2012. Diversity and origin of chinese troglobitic Carabid beetles (Coleoptera : Carabidae). Proceedings of the Guangxi International Symposium of Karst Cave Biodiversity Conservation, Nanning, China, 1-4.XII.2012. Pp. 6–8.
  • 32. Faille, A., Bourdeau, C., Fresneda, J. 2012. Molecular phylogeny of the Trechus brucki group, with description of two new species from the Pyreneo-Cantabrian area (France, Spain) (Coleoptera, Carabidae, Trechinae). Zookeys. 217: 11–51.
  • 31. Perreau, M., Faille, A. 2012. Contribution à la phylogénie et la systématique des Aleocharinae hypogés d'Afrique du nord (Coleoptera : Staphylinidae). Annales de la Société Entomologique de France (n.s.). 48(1-2): 81–96.


  • 30. Faille, A., Fresneda, J., Bourdeau, C. 2011. Les Molopina hypogés des Pyrénées avec la description d’une nouvelle espèce de Zariquieya Jeannel, 1924 d’Espagne (Coleoptera, Carabidae, Pterostichini). Zoosystema. 33(4): 429–441.
  • 29. Hendrich, L., Faille, A., Hawlitschek, O., Taenzler, R. 2011. Wiederfund des Schwimmkäfers Graphoderus bilineatus (DE GEER, 1774) (Coleoptera, Dytiscidae) nach über 25 Jahren in Bayern. Nachrichtenblatt der bayerischen Entomologen. 60(3/4) : 59–65.
  • 28. Faille, A., Bourdeau C. 2011. Une nouvelle espèce de Trechinae troglobie du versant sud des Pyrénées (Coleoptera, Carabidae, Trechinae). Bulletin de la Société Entomologique de France. 116(3): 261–267.
  • 27. Junger, B., Faille, A. 2011. Remarkable discovery in a cave of south west Morocco: Siagona taggadertensis n.sp. (Carabidae: Siagoninae). Annales de la Société Entomologique de France (n.s.). 47(1-2): 162–167.
  • 26. Fresneda, J., Bourdeau, C., Faille, A. 2011. Le genre Paratroglophyes Fourès, 1954 : description d’une nouvelle espèce du massif du Mont Valier, Pyrénées Centrales, France. (Coleoptera, Leiodidae, Cholevinae, Leptodirini). Bulletin de la Société Entomologique de France. 116(3): 355–363.
  • 25. Faille, A., Casale, A., Ribera, I. 2011. Phylogenetic relationships of west Mediterranean troglobitic Trechini groundbeetles (Coleoptera: Carabidae). Zoologica Scripta. 40(3) : 282–295.


  • 24. Fresneda, J., Bourdeau, C., Faille, A. 2010. Descripción de Bathysciola liqueana sp. n. de los Pirineos centrales (Francia). Designación de lectotipos y datos de distribución de las especies del grupo de B. meridionalis (Jacquelin du Val, 1854) (Insecta, Coleoptera, Leiodidae, Cholevinae, Leptodirini). Animal Biodiversity and Conservation. 33(2): 131–142.
  • 23. Ribera, I., Faille, A. 2010. A new microphthalmic stygobitic Graptodytes from Morocco, with a molecular phylogeny of the genus (Coleoptera, Dytiscidae). Zootaxa. 2641: 1–14.
  • 22. Faille, A. 2010. Les Coléoptères troglobies de Sulawesi (Indonésie). Description du mâle d’Eustra saripaensis Deuve (Coleoptera, Paussidae, Ozaeninae). Bulletin de la Société Entomologique de France. 115(3): 375–380.
  • 21. Fresneda, J., Bourdeau, C., Faille, A. 2010. Sobre la presencia de Catops subfuscus Kellner, 1846 en los Pirineos (Coleoptera, Leiodidae, Cholevinae, Catopini). Arxius de Miscel.lània Zoològica. 8: 9–14.
  • 20. Faille, A., Bourdeau, C., Fresneda, J. 2010. A new species of blind Trechinae from the Pyrenees of Huesca, and its position within Aphaenops (sensu stricto) (Coleoptera: Carabidae: Trechini). Zootaxa. 2566: 49–56.
  • 19. Fresneda, J., Fery, H., Faille, A. 2010. El complejo de Bathysciola ovata (Kiesenwetter, 1850): designación de lectotipos, establecimiento de sinonímias y consideraciones taxonómicas y corológicas. (Coleoptera, Leiodidae, Cholevinae, Leptodirini). Boletín de la Sociedad Entomológica Aragonesa. 46: 95–104.
  • 18. Faille, A., Bourdeau, C., Fresneda, J. 2010. Une seconde énigme biogéographique revisitée : Aphaenops (Cerbaphaenops) bourdeaui Coiffait, 1976 : (Coleoptera : Carabidae). Bulletin de la Société Entomologique de France. 115(2): 219–224.
  • 17. Faille, A., Ribera, I., Deharveng, L., Bourdeau, C., Garnery, L., Queinnec, E., Deuve, T. 2010. A molecular phylogeny shows the single origin of the Pyrenean subterranean Trechini ground beetles (Coleoptera: Carabidae). Molecular Phylogenetics and Evolution. 54: 97–105.


  • 16. Faille, A., Bourdeau, C., Fresneda, J. 2009. Eskualdunella delespierrei Coiffait, 1950: La clé inattendue d’une énigme biogéographique (Coleoptera: Leiodidae, Cholevinae, Leptodirini). Bulletin de la Société Entomologique de France. 114(4): 469–473.
  • 15. Santamaria, S., Faille, A. 2009. New species of Laboulbenia and Rhachomyces (Laboulbeniales, Ascomycota), some of them polymorphic, parasitic on termiticolous ground beetles from Tropical Africa. Nova Hedwigia. 89(1-2): 97–120.
  • 14. Fresneda, J., Bourdeau, C., Faille, A. 2009. El género Trapezodirus Jeannel, 1924: consideraciones biogeográficas y presencia en el medio hipogeo del macizo de Cotiella (Pirineos, Huesca, España) (Coleoptera, Leiodidae, Cholevinae, Leptodirini). Cotiella. 1: 24–30, 40.
  • 12. Fresneda, J., Bourdeau, C., Faille, A. 2009. Baronniesia delioti gen. n. sp. n., a new subterranean Leptodirini from the French Pyrenees (Coleoptera, Leiodidae, Cholevinae). Zootaxa. 1993:1–16.


  • 11. Faille, A. 2008. Estudis genètics de la fauna subterrània. Mètode de comparació de les seqüències d'ADN en coleòpters cavernícoles. espeleobloc.blogspot.com/2008/12/estudis-gentics-de-la-fauna-subterrnia.html
  • 10. Faille, A. 2008. Subterranean fauna of Pyrenees. Proceedings of the Ivth European Speological Congress, Vercors 2008. Spelunca Mémoires. N°33:60–64.
  • 9. Faille, A., Bourdeau C. 2008. Une nouvelle espèce de Trechinae endogé des Pyrénées Occidentales (Coleoptera : Carabidae). Bulletin de la Société Entomologique de France. 113(3): 307–311.


  • 8. Faille, A., Déliot, P., Queinnec, E. 2007. A new cryptic species of Aphaenops (Coleoptera, Trechinae) from french Pyrenean cave: Congruence between morphometrical and geographical data confirm species isolation. Annales de la Société Entomologique de France (n.s.). 43(3): 363–370.
  • 7. Santamaria, S., Faille, A. 2007. Rhachomyces (Ascomycota, Laboulbeniales) parasites on cave-inhabiting Carabidae beetles from Pyrenees. Nova Hedwigia. 85 (1-2):159–186.


  • 6. Faille, A. 2006. Endémisme et adaptation à la vie cavernicole chez les Trechinae pyrénéens (Coleoptera : Carabidae). Approches moléculaire et morphométrique. Ph.D. dissertation, Muséum National d’Histoire Naturelle, Paris, 319 p.
  • 5. Faille, A., Fresneda, J., Déliot, P., Bourdeau, C. 2006. Description du mâle d’Aphaenops catalonicus Escolà & Canció (Coleoptera, Trechinae). Bulletin de la Société Entomologique de France. 111(2): 247–250.
  • 4. Faille, A. 2006. Nouvelles données concernant les coléoptères Trechinae des gouffres d’altitude de la chaîne axiale pyrénéenne. Compte-Rendu Las Olas 2006. Spéléo Club du Comminges : 23–24.


  • 3. Bouyon, H., Faille, A. 2004. Compte-rendu de la sortie ACOREP organisée à Fontainebleau le 29 mars 2004 à l’occasion de l’abattage de trois vieux chênes aux Ventes à la Reine. Le Coléoptériste. 7(2) : 125–127.


  • 2. Faille, A. 2002. Le Platypsyllus castoris Rits., coléoptère ectoparasite du castor. Bulletin d’Information du Réseau de Correspondants CASTOR. Office National de la Chasse et de la Faune Sauvage. 13 : 13–15.
  • 1. Faille, A. 2002. Notes sur quelques raretés d’Ile-de-France. Le Coléoptériste. 5(2) : 121–123.



  • Curator Coleoptera, Staatliches Museum für Naturkunde Stuttgart


  • Contract Researcher, Institut de Biologia Evolutiva, Barcelona, ​​Spanien


  • DFG Research Fellow, Zoologische Staatssammlung München, Deutschland


  • Humboldt Research Fellow, Zoologische Staatssammlung München, Deutschland


  • Contract Researcher, Muséum National d'Histoire Naturelle, Paris, Frankreich


  • Postdoctoral Researcher, Institut de Biologia Evolutiva, Barcelona, ​​Spanien


  • Postdoctoral Researcher, Museo Nacional de Ciencias Naturales, Abteilung Biodiversität und Evolution, Madrid, Spanien



  • PhD in Systematics and Evolution (Zoology) (MNHN, Paris, France – summa cum laude)


  • Master in Zoological and Botanical Systematics (MNHN, Paris, France)


  • Master in Biology (LAVAL University, Québec, Canada / Université PARIS XI Orsay)

Funded Projects


  • Programa Estatal de Fomento de la Investigación Científica y Técnica de Excelencia, Spain - « CONEVOL : COnvergent EVOLution in subterranean Coleoptera »


  • German Research Foundation (DFG) - Munich, Germany - “Contraction of life cycle, Evolution and Diversification of cave beetles” (DFG FA 1042/1-1)


  • German Research Foundation (DFG) - Munich, Germany – « Evolution of subterranean ground beetles in the Western Palearctic » (DFG BA 2152/14-1)


  • Humboldt Research Fellowship - Alexander von Humboldt Foundation, Munich, Germany - « Molecular evolution and subterranean adaptations: Phylogeny and convergence in Alpine cave beetles »


  • National Center for Scientific Research (CNRS), Contract - UMR 7205 CNRS / MNHN, Paris, France - Partnership MNHN / CNRS / Syndicat      Intercommunal à Vocation Unique « LA VERNA » (SIVU), « Impact des Aménagements de la grotte de la Verna sur les populations d’insectes cavernicoles »


  • Consejo Superior de Investigaciones Científicas (CSIC), Contract - Institut de Biologia Evolutiva, Barcelona, Spain - «Molecular  Phylogeny of Pyrenean Trechinae»


  • Regional Directorate for Environment Midi-Pyrénées, Contract - PASTEL Funding « Cartography of Biodiversity : Endemic and relictual Invertebrates from Pyrenees »
  • Synthesys program - Museo Nacional de Ciencias Naturales, Madrid, Spain (ES-TAF-1540) « Endemism and adaptations to the subterranean life in a group of insects : the Aphaenops / Hydraphaenops species complex (Coleoptera, Carabidae, Trechinae) »


  • Legs Prévost Grant of the MNHN, Paris, France


  • Travel Grants (G. Cousin) – Société Entomologique de France


  • ABC grant (Access to Belgian Collections) - Royal Belgian Institute of Natural Sciences, Bruxelles, Belgium – « ESEM (Environmental Scanning Electron Microscope) Study of Leiodidae of RBINS collections »


 Algeria, Austria, Azerbaijan, Bosnia and Herzegovina, Bulgaria, Canary Islands, China, France, French Guyana, Georgia, Germany, Indonesia, Italia, Madagascar, Mallorca, Morocco, Romania, Sicily, Spain, Turkey, Vietnam

International expeditions

NAMOROKA 2016 - Biodiversity inventory of the Tsingy de Namoroka National Park, Madagascar , Madagaskar - MNHN-Expedition


LENGGURU 2014 - Exploration of biodiversity of the Karsts of Western Papua:


Biodiversity of the Annamitic chain (Vietnam) : endemism and altitudinal gradients 2013 - Mission Labex BCDIV, Hon-Ba-Massiv (Nha Trang)

WIN-TIMDOUINE 2008 - Subterranean exploration of the largest cave of Africa


SULAWESI 2007 - Zoological investigations in the karsts of South and Southeast Sulawesi