PD Dr. Mike Thiv

Profilbild Mike Thiv

Head of Botany department

Ferns and Flowering Plants (Pteridophyta and Phanerogamae)


+49 (0) 711 / 8936-205
+49 (0) 711 / 8936-100

Research interests

I am interested in plant systematics, biogeography (especially the Eritreo-Arabian region, the Mediterranean and Macaronesia) and floristics. My main taxonomic expertise is in Gentianaceae, Pulmonaria (Boraginaceae). I mainly participate in QT 2 and QT3.

Researchgate   Google Scholar




  • Krause, C., Oelschlägel, B., Mahfoud, H., Frank, D., Lecocq, G., Shuka, L., Neinhuis, C., Vargas, P., Tosunoglu, A., Thiv, M., Wanke, S. 2022: The evolution of the Aristolochia pallida complex (Aristolochiaceae) challenges traditional taxonomy and reflects large‐scale glacial refugia in the Mediterranean. Ecology and evolution 12 (4), e8765.


  • Thiv, M. Gouveia, M., de Sequeira, M.M. 2021: The Madeiran laurel forest endemic Goodyera macrophylla (Orchidaceae) is related to American orchids. Anales del Jardín Botánico de Madrid 78 (2), e116-e116.
  • Roth-Nebelsick, A., Thiv, M., Malkowsky, Y, Schott, R, Heyer A. 2021. Structure and functional anatomy of the gas exchange apparatus of leafless orchids: evidence for a control mechanism? Botanical Journal of the Linnean Society 197 (2), 249-262.


  • Borsch T., Stevens A.-D., Häffner E., Güntsch A., Berendsohn W.G., Appelhans M.S., Barilaro C., Beszteri B., Blattner F.R., Bossdorf O., Dalitz H., Dressler S., Duque-Thüs R., Esser H.-J., Franzke A., Goetze D., Grein M., Grünert U., Hellwig F., Hentschel J., Hörandl E., Janßen T., Jürgens N., Kadereit G., Karisch T., Koch M.A., Müller F., Müller J., Ober D., Porembski S., Poschlod P., Printzen C., Röser M., Sack P., Schlüter P., Schmidt M., Schnittler M., Scholler M., Schultz M., Seeber E., Simmel J., Stiller M., Thiv M., Thüs H., Tkach N., Triebel D., Warnke U., Weibulat T., Wesche K., Yurkov A., Zizka G. (2020): A complete digitization of German herbaria is possible, sensible and should be started now. – Research Ideas and Outcomes, 6: e50675. doi.org/10.3897/rio.6.e50675


  • Thiv, M., Reyes-Betancort, J.A., Fragman-Sapir, O. (2019): Lifeforms as criterion for species delimitation: Are Aristida adscensionis and A. coerulescens (Aristidoideae, Poaceae) two species? Phytotaxa 393 (1): 67-74.
  • Schüßler, C., Bräuchler, C., Reyes‐Betancort, J.A., Koch, M.A., Thiv, M. (2019): Island biogeography of the Macaronesian Gesnouinia and Mediterranean Soleirolia (Parietarieae, Urticaceae) with implications for the evolution of insular woodiness. Taxon 68 (3): 537-556.


  • Schüssler, C., Thiv, M., Wörz, A. (2018): Indigenous and synanthropic populations of Galanthus nivalis L. in Baden-Württemberg. Integrative Systematics, 1: 7–15.


  • Plenk, K., Bardy, K., Höhn, M., Thiv, M., Kropf, M.
    No obvious genetic erosion, but evident relict status at the westernmost range edge of the Pontic-Pannonian steppe plant Linum flavum L. (Linaceae) in Central Europe. Ecology and Evolution. 2017;1–13. DOI: 10.1002/ece3.2990.
  • Krause, C., Wörz, A., Thiv, M.
    Molecular analysis of the extinct Central European endemic Saxifraga oppositifolia subsp. amphibia and its implications on glaciation biogeography. Alpine Botany 127: 85-95. DOI 10.1007/s00035-016-0183-5.


  • Cuccuini, P., Nepi, C., Abuhadra, M. N., Banfi,  E., Domina, G, Luccioli, E., Miranda, S., Pagitz, K., Thiv, M., Vela, E.
    The Libyan Collections in FI (Herbarium Centrale Italicum and Webb Herbarium) and Studies on the Libyan Flora by R. Pampanini – Part 2. Fl. Medit. 26: 81-143. doi: 10.7320/FlMedit26.081.
  • Betzin, A., Thiv, M., Koch, M.A.
    Diversity hotspots of the laurel forest on Tenerife, Canary Islands: a phylogeographic study of Laurus and Ixanthus. Annals of Botany  118 (3): 495-510. doi:10.1093/aob/mcw124.
  • Thiv, M.
    Sphenostemonaceae & Vahliaceae. In Kadereit, J. W., Bittrich, V. (eds.): The Families and Genera of Vascular Plants: Eudicots. Vol. 14, pp. 363-366, 381-383. Springer Verlag, Hamburg.
  • Koch, M.A., Meyer, N., Engelhardt, M., Thiv, M., Bernhardt, K.-G., Michling, F.
    Morphological and genetic variation of highly endangered Bromus species and the status of these Neolithic weeds in Central Europe. Plant Syst. Evol. 302: 515. doi 10.1007/s00606-016-1279-5


  • Wörz, A, Thiv, M.
    The temporal dynamics of a regional flora—The effects of global and local impacts. Flora 217: 99-108.
  • Harter, D. E. V., Thiv, M., Weig, A., Jentsch, A., Beierkuhnlein, C.
    Spatial and ecological population genetic structures within two island-endemic Aeonium species of different niche width. Ecology and Evolution. 5(19): 4327–4344. doi: 10.1002/ece3.1682.
  • Kondraskov, P., Schütz, N., Schüßler, C., Menezes de Sequeira, M., Santos Guerra A., Caujapé-Castells, J., Jaén-Molina, R., Marrero-Rodríguez, A., Koch, M. A., Linder, P., Kovar-Eder, J., Thiv, M.
    Biogeography of Mediterranean Hotspot Biodiversity: Re-Evaluating the 'Tertiary Relict' Hypothesis of Macaronesian Laurel Forests. PLoS ONE 10(7): e0132091. doi:10.1371/journal.pone.0132091


  • Schneeweiss, G.M., Pachschwöll, C., Tribsch, A., Schönswetter, P., Barfuss, M. H. J., Esfeld, K., Weiss-Schneeweiss, H., Thiv, M.
    Molecular phylogenetic analyses identify Alpine differentiation and dysploid chromosome number changes as major forces for the evolution of the European endemic Phyteuma (Campanulaceae). Mol. Phylogenet. Evol. 69: 634-652.
  • Pleines, T., Esfeld, K., Blattner, F. R., Thiv, M.
    Ecotypes and genetic structure of Rhinanthus alectorolophus (Orobanchaceae) in southwestern Germany. Plant. Syst. Evol. 299: 1523-1535.


  • Liede-Schumann, S., Kong, H., Meve, U., Thiv, M. (2012): Vincetoxicum and Tylophora (Apocynaceae: Asclepiadoideae: Asclepiadeae) - two sides of the same medal: Independent shifts from tropical to temperate habitats. Taxon 61: 803–825.
  • Španiel, S., Marhold, K., Thiv, M., Zozomová-Lihová, J. (2012): A new circumscription of Alyssum montanum subsp. montanum and A. montanum subsp. gmelinii (Brassicaceae) in Central Europe: molecular and morphological evidence. Bot. J. Linn. Soc. 169: 378-402.
  • Wörz, A., Hohmann, N., Thiv, M. (2012): Morphological and molecular diversity of some populations of Gagea (Liliaceae) in Southwest Germany. Stuttgarter Beitr. Naturk. Ser. A Neue Serie 5: 1–11


  • Alejandro, G. J. D., Meve, U., Mouly, A., Thiv, M., Liede-Schumann, S. (2011): Molecular phylogeny and taxonomic revision of the Philippine endemic Villaria Rolfe (Rubiaceae). Pl. Syst. Evol. 296: 1-20.
  • Thiv, M., Niet, T v. d., Rutschmann, F., Thulin, M., Brune, T., Linder, H P. (2011): Trans-Atlantic and trans-African disjunctions of Thamnosma (Rutaceae): Intercontinental long distance dispersal and local differentiation in the arid biome. Am. J. Bot. 98(1): 76–87


  • Alejandro, G. J. D., Meve, U., Uy, M., Mouly, A., Thiv, M., Liede-Schumann, S. (2010): Molecular support of the classification of Greeniopsis Merr. in Aleisanthieae (Rubiaceae), with a revision of the genus. Taxon 59(5): 1547–1564.
  • Thiv, M., Esfeld, K., Koch, M. (2010): Studying adaptive radiation at the molecular level: a case study in the Macaronesian Crassulaceae-Sempervivoideae. In Glaubrecht, M. (ed.) Evolution in Action - Adaptive Radiations and the Origins of Biodiversity. Pp. 35-59. Springer Verlag, Heidelberg.
  • Thiv, M., Thulin M., Hjertson M., Kropf, M., Linder H.P. (2010): Evidence for a vicariant origin of Macaronesian-Eritreo/Arabian disjunctions in Campylanthus Roth (Plantaginaceae). Mol. Phylogenet. Evol. 54: 606-616.


  • Thiv, M., Ghogue,  J.- P., Grob, V., Huber,  K., Pfeifer, E., Rutishauser, R. (2009): How to get off the mismatch at the generic rank in African Podostemaceae? Pl. Syst. Evol. 283: 57-77.
  • Esfeld, K, Niet, T. van der, Seifan, M., Koch, M., Thiv, M. (2009): Little interspecific pollen transfer despite overlap in pollinators between sympatric Aeonium (Crassulaceae) species pairs. Flora 204: 709-717.
  • Pfeifer, E., Grob, V., Thiv, M., Rutishauser, R. (2009): Stonesia ghoguei. Comparative morphology of a new Cameroonian species (Podostemaceae, Podostemoideae). Novon 19: 102-116.


  • Wörz, A., Engelhardt, M., Hölzer, A., M. Thiv (2008): Aktuelle Verbreitungskarten der Farn-  und Blütenpflanzen Baden-Württembergs. http://www.flora.naturkundemuseum-bw.de
  • Brune T., Thiv, M., Haas, K. (2008): Equisetum (Equisetaceae) species or hybrids? ISSR fingerprinting profiles help improve diagnoses based on morphology and anatomy. Pl. Syst. Evol. 274: 67-81.


  • Moline, P., Thiv, M., Ameka, G.K., Ghogue, J.-P., Pfeiffer, E., Rutishauser, R. (2007): Comparative Morphology and Molecular Systematics of African Podostemaceae-Podostemoideae, with Emphasis on Dicraeanthus and Ledermanniella from Cameroon. Int. J. Plant Sci. 168: 159-180.
  • Thiv, M., Meve, U. (2007): A phylogenetic study of Echidnopsis Hook. f. (Apocynaceae-Asclepiadoideae) - taxonomic implications and the colonization of the Socotran archipelago. Pl. Syst. Evol. 265: 71-86.


  • Thiv, M., Thulin M., Kilian, N., Linder, H.P. (2006): Eritreo-Arabian affinities of the Socotran flora as revealed from the molecular phylogeny of Aerva Forssk. (Amaranthaceae). Syst. Bot. 31(3): 560-570.


  • Hoechstetter, S., Wahr, J., Knapp, S., Klotz, S., Uhl, D., Thiv, M., Mosbrugger, V. (2005): Biodiversity patterns of vascular plants in Baden-Württemberg (SW Germany) - a geostatistical analysis of abiotic factors. Stuttgarter Beitr. Naturk. Ser. A 676: 1-25.


  • Thiv, M. (2004): Carlemanniaceae. In Kadereit, J. W. (ed.), The Families and Genera of Vascular Plants: Euasterids. Springer Verlag, Hamburg.


  • Thiv, M. (2003): A taxonomic revision of Canscora, Cracosna, Duplipetala, Hoppea, Microrphium, Phyllocyclus, and Schinziella (Gentianaceae-Canscorinae). Blumea 48: 1-46.


  • Thiv, M., Kadereit, J. W. (2002): A morphological-cladistic analysis of Gentianaceae- Canscorinae and the evolution of anisomorphic androecia in the subtribe. Syst. Bot. 27(4): 780-788.
  • Thiv, M., Struwe, L., Albert, V. A., Kadereit, J. W.: Tribe Chironieae. Pp. 108-137. In: Struwe, L, Hagen, K. B. von, Kadereit, J. W., Klackenberg, J., Nilsson, J. S., Thiv, M., Albert, V. A. (2002): Systematics, character evolution, and biogeography of Gentianaceae, including a new tribal and subtribal classification. In: Struwe, L., Albert, V. A. (eds.) Gentianaceae - Systematics and Natural History. Cambridge University Press, Cambridge.
  • Thiv, M., Kadereit, J. W.: Tribe Saccifolieae. Pp. 55-66. In: Struwe, L, Hagen, K. B. von, Kadereit, J. W., Klackenberg, J., Nilsson, J. S., Thiv, M., Albert, V. A. (2002): Systematics, character evolution, and biogeography of Gentianaceae, including a new tribal and subtribal classification. In: Struwe, L., Albert, V. A. (eds.), Gentianaceae - Systematics and Natural History. Cambridge University Press, Cambridge.
  • Thiv, M. (2002): Gentianaceae. In: W. Greuter, Rodriguez, R. R., Manitz, H. (eds.) Flora de la Republica de Cuba. Vol. 6: 1-40. Koeltz, Königstein.


  • Thiv, M. (2000): Molekulare und morphologische Phylogenie und Biogeographie der Gentianaceae-Chironieae und eine Revision der Gattungen Canscora, Cracosna, Duplipetala, Hoppea, Microrphium, Phyllocyclus und Schinziella (Gentianaceae-Canscorinae). Tectum Verlag Marburg.


  • Thiv, M., Struwe, L., Kadereit, J. W. (1999): The phylogenetic relationships and evolution of the Canarian laurel forest endemic Ixanthus viscosus (Ait.) Griseb. (Gentianaceae): evidence from matK and ITS sequence variation, and floral morphology and anatomy. Pl. Syst. Evol. 218 (3-4): 299-317.
  • Thiv, M., Struwe, L., Albert, V. A., Kadereit, J. W. (1999): The phylogenetic relationships of Saccifolium bandeirae Maguire & Pires (Gentianaceae) reconsidered. Harvard Pap. Bot 4: 519-526.


  • Struwe, L., Thiv, M., Kadereit, J. W., Pepper, A. S.-R., Motley, T. J., White, P. J., Rova, J. H. E., Potgieter, K., Albert, V. A. (1998): Saccifolium (Saccifoliaceae), an endemic of Sierra de La Neblina on the Brazilian-Venezuelan frontier, is related to a temperate-alpine lineage of Gentianaceae. Harvard Pap. Bot. 3: 199-214.


  • Sauer, W., Kleinsteuber, A., Thiv, M. (1996): Pulmonaria. In Sebald, O, Seybold, S., Philippi, G., Wörz, A. (eds.) Die Farn- und Blütenpflanzen Baden-Württembergs. Vol. 5: 90-98, Ulmer Verlag Stuttgart.


  • Sauer, W., Thiv, M. (1995): Die Gattung Pulmonaria (Boraginaceae) in Baden-Württemberg. Jh. Ges. Naturkde. Württemberg 151: 281-344.



  • Habilitation at University of Heidelberg

since 2004

  • Head of Botany Department at Natural History Museum Stuttgart, Germany

since 2003

  • Curator at Natural History Museum Stuttgart, Herbarium STU, Germany


  • Postdoc at the Institute of Systematic Botany, University of Zurich, Switzerland; Project "Molecular phytogeography of selected plant taxa of the Socotra Archipelago (Yemen)" with Prof. H. P. Linder


  • Curatorial work (internship) at Botanic Garden and Botanic Museum Berlin-Dahlem, Germany


  • Research Scientist at the Institut für Spezielle Botanik, Johannes Gutenberg-Universität Mainz, Germany



  • Dissertation (Prof. J. W. Kadereit): "Molecular and morphological phylogeny and biogeography of Gentianaceae-Chironieae and a revision of Canscora, Cracosna, Duplipetala, Hoppea, Microrphium, Phyllocyclus, and Schinziella (Gentianaceae-Canscorinae)"


  • Diploma thesis (Prof. W. Sauer): "Caryological and biogeographic studies on Pulmonaria L. (Boraginaceae) in Southwest-Germany"


  • Study of Biology, Eberhard-Karls-Universität Tübingen, Germany


  • As PI ca. 800.000 €  Funding

Editorial Board

  • Associate editor: Plant Systematics and Evolution, Plant and Fungal Systematics, PeerJ, Philippine Journal of Systematic Biology