Dr. Arnold H. Staniczek

Profilbild Arnold Staniczek

Senior Researcher and Curator

Ephemeroptera, Odonata, Plecoptera, Trichoptera


+49 (0) 711 / 8936-239
+49 (0) 711 / 8936-100

Research interests

My research program is centered on the basal groups of winged insects. It focuses on taxonomy and systematics of extant and fossil mayflies and stoneflies as well as other fossil groups of Palaeoptera. Comparative studies on the functional morphology of mayfly nymphs are closely linked to the question of their phylogenetic positions. Local faunistic studies contribute to a reference collection of water insects of the Federal State of Baden-Württemberg.

Researchgate   Academia   ORCID



in preparation

  • Staniczek, A.H., Bergmann, P., Van de Kamp, T. & Weide, D.  (in prep.): A wolf among sheep: Larval head morphology of the carnivorous mayfly Ameletopsis perscitus (Ephemeroptera: Ameletopsidae). 
  • Gamba, R., Van de Kamp, T., Staniczek, A.H. (in prep.): When mayflies have an erection: Functional morphology of genitalia in Ecdyonurus venosus (Insecta: Ephemeroptera: Heptageniidae) during copulation. Zoological Journal of the Linnean Society.
  • Staniczek, A.H., Sroka, P., Bojková, J., Nejat, F., Namin, I.J., Abdoli, A., Godunko, R.J. (in prep.): A new species of Prosopistoma (Insecta, Ephemeroptera, Prosopistomatidae) from Iran, with key to Western Palearctic species. Zookeys.
  • Staniczek, A.H., Schmalfuss, H. (in prep.): In Memoriam: Gerhard Mickoleit. Integrative Systematics: Stuttgart Contributions to Natural History. 




  • Domínguez, E., van de Kamp, T., Mikó, I., Cuezzo, M.G., Staniczek, A.H. (2023):The function of wing bullae in mayflies (Insecta: Ephemeroptera) opens new insights into the early evolution of Pterygota. BMC Biology 21(268): 1-23.
  • Storari, A.P., Staniczek, A.H., Godunko, R.J. (2023): Author Correction: A new Gondwanan mayfly family from the Lower Cretaceous Crato Formation, Brazil (Ephemeroptera: Siphlonuroidea: Astraeopteridae fam. nov.). Scientific Reports 13 (19274): 1-2. 
  • ​​​​​​Storari, A.P., Staniczek, A.H., Godunko, R.J. (2023): A new Gondwanan mayfly family from the Lower Cretaceous Crato Formation, Brazil (Ephemeroptera: Siphlonuroidea: Astraeopteridae fam. nov.). Scientific Reports 13 (11735): 1-14. 


  • Prevec, R., Nel, A., Day, M.O., Muir, R., Matiwane, A., Kirkaldy, A.P., Moyo, S., Staniczek, A.H., Cariglino, B., Maseko, Z., Kom, N., Rubidge, B.S., Garrouste, R., Holland, A., Barber-James, H.M. (2022): South African Lagerstätte reveals middle Permian Gondwanan lakeshore ecosystem in exquisite detail. Communications Biology 5: 1154, 1-14. https://doi.org/10.1038/s42003-022-04132-y
  • Godunko, R.J., Alba-Tercedor, J., Grabowski, M., Rewicz, T., Staniczek, A.H. (2022) Cenozoic origins of the genus Calliarcys (Insecta, Ephemeroptera) revealed by Micro-CT, with DNA barcode gap analysis of Leptophlebiinae and Habrophlebiinae. Scientific Reports 12 (15228): 1-17. https://doi.org/10.1038/s41598-022-18234-4
  • Staniczek, A.H., Storari, A.P., Godunko, R.J. (2022): Revised systematics, phylogeny, and paleontology of the mayfly family Baetiscidae (Insecta: Ephemeroptera). Arthropod Systematics and Phylogeny 80: 389-409. DOI: 10.3897/asp.80.e82845.
  • Sroka, P., Staniczek, A.H. (2022):  Evolution of filter-feeding in aquatic insects dates back to the Middle Triassic: New evidence from stemgroup mayflies (Insecta: Ephemerida) of Grès à Voltzia, Vosges, France. Papers in Palaeontology 8(4): e1456, 1-17. https://doi.org/10.1002/spp2.1456
  • Van de Kamp, T., Mikó, I., Staniczek, A.H., Eggs, B., Bajerlein, D., Faragó, T., Hagelstein, L., Hamann, E., Pretzsch, R., Baumbach, T., Janšta, P., Krogmann, L. (2022): Evolution of flexible biting in hyperdiverse parasitoid wasps. Proceedings of the Royal Society B (Biological Sciences). https://doi.org/10.1098/rspb.2021.2086


  • Schawaller, W., Staniczek, A.H. (2021): In Memoriam: Roland Grimm. Integrative Systematics: Stuttgart Contributions to Natural History 4(1): 1-5.

  • Storari, A.P.,  Godunko,R.J., Salles, F.F., Saraiva, A.A.F., Staniczek, A.H., Rodrigues, T. (2021): An overview of the Hexagenitidae (Ephemeroptera) from the Crato Formation (Aptian, Lower Cretaceous) of Brazil, with the description of a new species. Historical Biology: 1-10.

  • Godunko, R.J., Martynov, A.V., Staniczek, A.H.(2021): First fossil record of the mayfly family Vietnamellidae (Insecta, Ephemeroptera) from Burmese Amber confirms its Oriental origin and gives new insights into its evolution. Zookeys 1036: 99-120.


  • Hrivniak, L., Sroka, P., Bojková, J., Godunko, R.J., Namin, I.J., Bagheri, S., Nejat, F., Abdoli, A., Staniczek, A.H. (2020): Diversity and distribution of Epeorus (Caucasiron) (Ephemeroptera: Heptageniidae) in Iran, with descriptions of three new species. Zookeys 947: 71-102.
  • Staniczek, A.H., Malzacher, P.,  Bojková, J., Sroka, P., Soldán, T., Namin, I.J., Nejat, F., Abdoli, A., Godunko, R.J. (2020): Caenidae (Insecta: Ephemeroptera) of Iran, with new records and re-description of the nymph of Caenis kopetdagi Kluge, 1985. Aquatic Insects 41(2): 106-130.
  • Hrivniak, L., Sroka, P., Godunko, R.J., Bojková, J., Soldán, T.,  Staniczek, A.H. (2020): The impact of Miocene orogenesis in diversification and biogeography of Caucasian Epeorus (Caucasiron) mayflies (Ephemeroptera: Heptageniidae). Molecular Phylogenetics and Evolution 146 (106735): 1-12.
  • Sroka, P. & Staniczek. A.H. (2020): Retention of cervical and abdominal gills in the adult of a new fossil stonefly (Insecta, Plecoptera, Petroperlidae) from mid-Cretacous Burmese amber. Cretaceous Research 107 (104277): 1-6.
  • Staniczek, A.H. (2020): Die diesjährige Massenemergenz der Eintagsfliege Ephoron virgo (Ephemeroptera: Polymitarcyidae) in Vaihingen/Enz, mit Anmerkungen zur aktuellen und historischen Verbreitung des Uferaases in Baden-Württemberg - Mitteilungen des Entomologischen Vereins Stuttgart 55: 3-13.



  • Godunko, R.J., Neumann, C., Staniczek, A.H. (2019): Revision of fossil Metretopodidae (Insecta: Ephemeroptera) in Baltic amber - Part 4: Description of two new species of Siphloplecton Clemens, 1915, with notes on the new S. jaegeri species group and with key to fossil male adults of SiphloplectonZookeys 898: 1-26.
  • Makarkin, V.N., Staniczek, A.H. (2019): A large-sized new genus of Babinskaiidae (Neuroptera: Myrmeleontoidea: Nymphidoidae) from mid-Cretaceous Burmese amber. Cretaceous Research 104: 104196.
  • Sroka, P., Bojková, J., Godunko, R.J., Soldán, T., Namin, I.J., Nejat, F., Abdoli,A., Staniczek, A.H. (2019): New Oligoneuriidae (Insecta: Ephemeroptera) from Iran. Zookeys 872: 101-126.


  • Malzacher, P., Staniczek, A.H. (2018): A new mayfly genus of the subfamily Caeninae (Ephemeroptera: Caenidae) from Sumatra and two new Caenis species from Papua New Guinea. Integrative Systematics 1: 35-45.
  • Sroka, P., Staniczek. A.H. , Kondratieff, B.C. (2018): "Rolling" Stoneflies (Insecta: Plecoptera) from Mid-Cretaceous Burmese amber. Peer J 6:e5354.
  • Bojková, J., Sroka, P., Soldán, T., Namin, J.I., Staniczek, A.H., Polášek, M., Hrivniak, L., Abdoli, A., Godunko, R.J. (2018): Initial commented checklist of Iranian mayflies (Insecta: Ephemeroptera), with new area records and description of Procloeon caspicum sp. n. (Baetidae). Zookeys 749: 87-123. 
  • Merker, S., Staniczek, A.H. (2018): The Zoological Collections of the Stuttgart State Museum of Natural History. In: Zoological Collections of Germany – The animal kingdom in its amazing plenty at museums and universities: 621-645. Springer, Heidelberg. pdf


  • Staniczek A.H., Godunko R.J. & Kluge N.J. (2017): Fossil record of the mayfly family Ephemerellidae (Insecta, Ephemeroptera), with description of a new species and first report of Ephemerellinae from Baltic Amber. Journal of Systematic Palaeontology 16 (15), 2018: 1319-1335.
  • Wartenberg, N. Reinhard, S., Nöllert, A., Staniczek, A.H. & Kupfer, A. (2017): Caddisfly larvae (Trichoptera: Phryganeidae) as scavengers of frog carcasses. Salamandra 53(3): 458-460.
  • Staniczek, A.H., Godunko, R.J., Krzeminski, W. (2017): A new fossil mayfly species of the genus Borinquena Traver, 1938 (Insecta: Ephemeroptera: Leptophlebiidae: Atalophlebiinae) from Miocene Dominican amber. Annales Zoologici 67(1): 113-119.
  • Godunko, R.J., Soldán, T., Staniczek, A.H. (2017): Baetis (Baetiscypronyx sp. nov., a new species of the Baetis alpinus species-group (Insecta: Ephemeroptera: Baetidae) from Cyprus, with annotated checklist of Baetidae in the Mediterranean islands. Zookeys 664: 1-32.


  • Malzacher, P., Staniczek, A.H. (2016): Two new genera of Caeninae (Ephemeroptera: Caenidae), with a cladistic analysis of the tribe Clypeocaenini. Stuttgarter Beiträge zur Naturkunde A, Neue Serie 9: 41-62.
  • Staniczek, A.H., Godunko, R.J. (2016): Revision of fossil Metretopodidae (Insecta: Ephemeroptera) in Baltic amber – Part 3: Description of two new species of Siphloplecton Clemens, 1915, with notes on the re-discovered lectotype of Siphloplecton macrops (Pictet-Baraban & Hagen, 1856). Zootaxa 4103(1): 1-24. 
  • Huang, D.-Y., Bechly, G., Nel, P., Engel, M.S., Prokop, J., Azar, D., Cai, C.-Y., van de Kamp, T., Staniczek, A.H., Garrouste, R., Krogmann, L., dos Santos Rolo, T., Baumbach, T., Ohlhoff, R., Shmakov, A.S., Bourgoin, T. & Nel, A. (2016): New fossil insect order Permopsocida elucidates major radiation and evolution of suction feeding in hemimetabolous insects (Hexapoda: Acercaria). Scientific Report 6, 23004: 1-9. (DOI: 10.1038/srep23004)


  • Sroka, P., Staniczek, A.H., Bechly, G. (2014): Revision of the giant pterygote insect Bojophlebia prokopi Kukalová-Peck, 1985 (Insecta: Hydropalaeoptera) from the Carboniferous of the Czech Republic, with a first cladistic study of fossil palaeopterous insects. Journal of Systematic Palaeontology 13(11): 963-982.
  • Staniczek, A.H., Sroka, P., Bechly, G. (2014): Neither silverfish nor fowl: The enigmatic Carboniferous Carbotriplura kukalovaeKluge, 1996 (Insecta: Carbotriplurida) is the putative fossil sister group of winged insects (Insecta: Pterygota). Systematic Entomology 39(4): 619-632.
  • Staniczek, A.H., Godunko, R.J. (2014): Revision of fossil Metretopodidae (Insecta: Ephemeroptera) in Baltic amber. - Part 2: Description of a new species of Metretopus Eaton, 1901. - Historical Biology 27(7): 861–867. 



  • Staniczek, A.H., Godunko, R.J. (2012): Revision of fossil Metretopodidae (Insecta: Ephemeroptera) in Baltic amber. - Part 1: Type specimens of hitherto described fossil species of Siphloplecton Clemens, 1915 and Metretopus Eaton, 1901, with description of four new fossil species of Siphloplecton. - Palaeodiversity 5: 57-87.




  • Staniczek, A. H. (ed.) (2009): International Perspectives in Mayfly and Stonefly Research. Proceedings of the 12th International Conference on Ephemeroptera and the 16th International Symposium on Plecoptera, Stuttgart 2008. - Aquatic Insects 31, Supplement 1: 1-747.  
  • Pescador, M. L., Richard, B. A., Hubbard, M. D., Staniczek, A. H. (2009): Evolution of Baetiscidae (Ephemeroptera): current state of knowledge of the family. - Aquatic Insects 31, Supplement 1: 137-147. 
  • Ober, S. V., Staniczek, A. H. (2009): A new genus and species of coenagrionid damselflies (Insecta, Odonata, Zygoptera, Coenagrionidae) from Vanuatu. - Zoosystema 31 (3): 485-497. 
  • Ogden, T. H., Gattolliat, J.-L., Sartori, M., Staniczek, A. H., Soldan, T., Whiting, M. F. (2009): Towards a new paradigm in mayfly phylogeny (Ephemeroptera): combined analysis of morphological and molecular data. - Systematic Entomology 34(4): 616-634. 


  • Staniczek, A. H. (2007): Ephemeroptera: mayflies. In: Martill, D., Bechly, G. & Heads, S. (eds). The Crato Fossil Beds of Brazil: Window into an Ancient World: 163-184. Cambridge (Cambridge University Press).   
  • Staniczek, A. H., Bechly, G. (2007): "Apterygota": Primarily wingless insects. In: Martill, D., Bechly, G. & Heads, S. (eds). The Crato Fossil Beds of Brazil: Window into an Ancient World: 149-154. Cambridge (Cambridge University Press).   
  • Malzacher, P., Staniczek, A. H. (2007): Caenis vanuatensis, a new species of mayflies (Ephemeroptera: Caenidae) from Vanuatu. - Aquatic Insects 29(4): 285-295. 
  • Misof B., Niehuis O., Bischoff I., Rickert A., Erpenbeck D., Staniczek A. H. (2007): Towards a 18S phylogeny of hexapods: Accounting for group specific character covariance in optimized mixed nucleotide/doublet models. - Zoology (Jena) 110(5): 409-429. 


  • Malzacher, P., Staniczek, A. H. (2006): Revision of the Madecocercinae (Ephemeroptera: Caenidae). - Aquatic Insects 28(3): 165-193. 
  • Misof, B., Niehuis, O., Bischoff, I., Rickert, A., Erpenbeck, D., Staniczek A. H. (2006): A hexapod nuclear SSU rRNA secondary structure model and catalogue of taxon-specific structural variation. - Journal of Experimental Zoology Part B: Molecular and Developmental Evolution 306B: 70-88.  


  • Staniczek, A. H. (2003): New fossil mayflies from Dominican amber (Insecta: Ephemeroptera: Leptophlebiidae: Atalophlebiinae) - Stuttgarter Beitr. Naturk. (B) 341: 1-22.
  • Staniczek, A. H. (2003): Capnopsis schilleri (Plecoptera: Capniidae) in Baden-Württemberg - Mitt. ent. V. Stuttgart 38(1): 9-12.  
  • Hitchings, T. R. & Staniczek, A. H. (2003): Nesameletidae (Insecta: Ephemeroptera). Fauna of New Zealand 46: 1-72. Manaaki Whenua Press, Lincoln, New Zealand.
  • Staniczek, A. H. (2003): Die Mandibel der dicondylen Insekten: Neue Erkenntnisse zur Klärung der basalen Spaltungsereignisse in der Phylogenie der Fluginsekten. Verh. Westd. Entomologentag 2001: 89-91, Löbbecke-Mus., Düsseldorf.    


  • Staniczek, A. H. & Bechly, G. (2002): First fossil record of the Nearctic mayfly family Baetiscidae in Baltic Amber (Insecta: Ephemeroptera). Stuttgarter Beitr. Naturk. (B), Nr. 322:1-11.  



  • Staniczek, A. H. (2000): The Mandible of Silverfish (Insecta:Zygentoma) and Mayflies (Ephemeroptera) – Its Morphology and Phylogenetic Significance. Zool. Anz. 239 (2000): 147-178. 


  • Staniczek, A. H. (1997): The Morphology of Siphlaenigma janae Penniket, 1962 (Ephemeroptera, Siphlaenigmatidae), and Its Significance for the Ground Plan of Baetoidea. - In: Landolt, P. & Sartori, M. (eds.): Ephemeroptera & Plecoptera: Biology-Ecology-Systematics. MTL, Fribourg: 536-549. 


  • Staniczek, A. H. (1996): The Mandibular Articulation of Mayflies (Insecta: Ephemeroptera) and Its Implications for the Phylogeny of the Basic Pterygote Lineages. Proc. XXth Int. Congr. Entomology: 2. 


  • Staniczek, A. H. (1994): Untersuchung zur postembryonalen Ontogenese von Baetis rhodani (Pictet, 1894) (Insecta: Ephemeroptera) unter besonderer Berücksichtigung der larvalen Abdominalmuskulatur und der Tracheenkiemen:1-80. Diplomarbeit der Eberhard-Karls-Universität Tübingen.

Popular science


  • Staniczek, A. H. (2022): Am Anfang war das Feuer, am Ende der Lichtsmog. S. 31-34 in: Petra Härtl: Anthropozän. Zeitalter? Zeitenwende? Zukunft? - Stuttgarter Beiträge zur Naturkunde, Serie C, 86.
  • Staniczek, A. H. (2022): Das Schicksal unserer Flüsse. S. 39-41 in: Petra Härtl: Anthropozän. Zeitalter? Zeitenwende? Zukunft? - Stuttgarter Beiträge zur Naturkunde, Serie C, 86.


  • Staniczek, A. H. (2021): Bernsteinkabinett. S. 138-157 in: Kovar-Eder et al.: Zeitreise durch die Erdgeschichte im Museum am Löwentor. - Stuttgarter Beiträge zur Naturkunde, Serie C, 85.


Staniczek, A. H. (2020): One fly does not make a summer: The mass flight of mayflies on the River Enz. - SMNS Science Blog: Understanding Nature.


  • Staniczek, A. H. (2019): Leben im Bernsteinwald. - Stuttgarter Beiträge zur Naturkunde, Serie C, 84: 1-160.



  • Staniczek, A. H. (2014): Fossiles Bindeglied deutet auf passive Gleitflieger an der Basis der geflügelten Insekten. - Naturwissenschaftliche Rundschau 67 (7): 368-369.
  • Staniczek, A. H. (2014): Evolution der Fluginsekten - Das 309 Millionen Jahre alte Fossil Carbotriplura kukalovae ist der nächste Verwandte der geflügelten Insekten. SMNS Science Blog: Understanding Nature.
  • Staniczek, A. H. (2014): Ein Jahr am Goldersbach - Langzeituntersuchung eines einheimischen Fließgewässers. - SMNS Science Blog: Understanding Nature.


  • Staniczek, A. H., Schmid, U., Mika, T. (2012): Sex - Motor der Evolution. - Stuttgarter Beiträge zur Naturkunde, Serie C, 72: 1-112.


  • Staniczek, A. H. (2007): Der Floh im Nasenloch: Parasitismus bei Papageien. - Naturwissenschaftliche Rundschau 60 (9): 479-480. 


  • Staniczek, A. H. (2005): Ein Fächerflügler aus dem Baltischen Bernstein - Naturwissenschaftliche Rundschau 58 (5): 271-272. 


  • Staniczek, A. H. (2003): Sind die Flügel bei Stabheuschrecken mehrfach neu entstanden? - Naturwissenschaftliche Rundschau 56 (7): 386-387. 
  • Staniczek, A. H. (2003): Eintagsfliegen - Manna der Flüsse. - Stuttgarter Beiträge zur Naturkunde, Serie C, 53: 1-80.  
  • Staniczek, A. H. (2003): Evolution der Wasserkäfer. - Naturwissenschaftliche Rundschau 56 (6): 331-333. 


  • Staniczek, A. H. (2002): Fossile Eintagsfliegen - Einblicke in die Welt urtümlicher Fluginsekten. Fossilien 19: 297-302.



  • Staniczek, A. H. (2013): Michael Schmitt: From Taxonomy to Phylogenetics - Life and Work of Willi Hennig. - Naturwissenschaftliche Rundschau 66(6): 325-326.


  • Staniczek, A. H. (2008): S. Coscaron and C.L. Coscarron Arias: Neotropical Simuliidae. A. Borkent and G.R. Spinelli: Neotropical Ceratopogonidae. Aquatic Biodiversity in Latin America, vols. 3 and 4. Pensoft (2007). - Aquatic Insects 30(1): 79-80.  
  • Staniczek, A. H. (2008): P.H. Langton & L.C.V. Pinder: Keys to the adult male Chironomidae of Great Britain and Ireland. FBA Scientific Publication 64 (2007). - Aquatic Insects 30(1): 79. 


  • Staniczek, A. H. (2007): M.J. Winterbourn et al.: Guide to the Aquatic Insects of New Zealand (4th ed.). Bulletin of the Entomological Society of New Zealand 14 (2006).  - Aquatic Insects 29(1): 76. 
  • Staniczek, A. H. (2007): E. Dominguez et al.: Ephemeroptera of South America. Aquatic Biodiversity in Latin America, vol.2. Pensoft (2006). - Aquatic Insects 29(1): 71-72. 


  • Staniczek, A. H. (2006): H. Malicky: Atlas of European Trichoptera (2nd ed.). Springer (2004). - Aquatic Insects 28(2): 163-164.
  • Staniczek, A. H. (2006): A. Tanida & A. Rossiter: Proceedings of the 11th International Symposium on Trichoptera. Tokai University Press (2005). - Aquatic Insects 28(2): 163.  


  • Staniczek, A. H. (2005):  R.S. Wilson & L.P. Ruse: A guide to the identification of genera of chironomid pupal exuvia occuring in Britain and Ireland. FBA Special Publication No.13 (2005). - Aquatic Insects 27(4): 310-311. 
  • Staniczek, A. H. (2005): Glenn B. Wiggins: Caddisflies: Underwater Architects. University of Toronto Press (2004). - Aquatic Insects 27(4): 309-310.. 
  • Staniczek, A. H. (2005): A.N. Nilsson & B.J. van Vondel: World Catalogue of Insects. Volume 7. Amphizoidae, Aspidytidae, Haliplidae, Noteridae and Paelobiidae (Coleoptera, Adephaga). Apollo Books (2005). - Aquatic Insects 27(4): 309. 
  • Staniczek, A. H. (2005): Stefan Richter, Walter Sudhaus: Kontroversen in der Phylogenetischen Systematik der Metazoa. - Naturwissenschaftliche Rundschau 58 (7): 397-398. 
  • Staniczek, A. H. (2005): Nikita Kluge: The Phylogenetic System of Ephemeroptera. Kluwer Academic Publishers (2004). - Aquatic Insects 27(1): 71-73. 


  • Staniczek, A. H. (2004): Gerhard Mickoleit: Phylogenetische Systematik der Wirbeltiere. - Naturwissenschaftliche Rundschau 57 (10): 579-580. 


  • Staniczek, A. H. (2003): Die Eulenfalter (Noctuidae) Mitteleuropas – ein interaktives Bestimmungswerk auf CD-ROM. Naturwissenschaftliche Rundschau 56 (2): 3. 


  • Staniczek, A. H. (2001): Atlas de Distribution des Ephemeres de Suisse (Insecta: Ephemeroptera). Fauna Helvetica 3 – by M. Sartori & P. Landolt. Entomologist's Monthly Magazine.



  • Kommissarische Leitung der Abteilung Entomologie am SMNS

seit 2003

  • Kurator für aquatische Insekten am Staatlichen Museum für Naturkunde Stuttgart


  • Wissenschaftlicher Volontär in der Abteilung Paläontologie am Staatlichen Museum für Naturkunde Stuttgart.


  • Promotion bei Dr. G. Mickoleit, Eberhard-Karls-Universität Tübingen: Der Larvenkopf von Oniscigaster wakefieldi McLachlan, 1873 (Insecta: Ephemeroptera: Oniscigastridae)


  • Studium der Biologie an der Eberhard-Karls-Universität Tübingen mit Auslandssemester an den Universitäten Helsinki und Oulu, Finnland.
  • Diplomprüfung in den Fächern Zoologie, Paläontologie, Genetik und Pflanzenphysiologie.
  • Diplomarbeit bei Dr. G. Mickoleit, Eberhard-Karls-Universität Tübingen: Die postembryonale Entwicklung der Tracheenkiemen bei der Larve von Baetis rhodani (Ephemeroptera: Baetidae)

Cooperation in scientific organisations


  • Mitglied im Vorstand der Gesellschaft zur Förderung des Naturkundemuseums Stuttgart e.V.


  • Mitglied im Ausschuss der Gesellschaft zur Förderung des Naturkundemuseums Stuttgart e.V.

seit 2009

  • Mitglied des "Permanent Committee of the International Conferences on Ephemeroptera"

Organised scientific congresses


  • Co-Organisation des Symposiums "Aquatic Insects", IXth European Conference of Entomology, 22.-27.8.2010, Budapest


  • Organisation des "International Joint Meeting on Ephemeroptera and Plecoptera 2008: XIIth International Conference on Ephemeroptera and XVIth International Symposium on Plecoptera, 8.-14.6.2008, Stuttgart"

Research stays

  • Naturhistorisches Museum, Vienna, Austria
  • Natural History Museum, London, UK
  • Muséum National d'Histoire Naturelle, Paris, France
  • Musée Cantonal de Zoologie, Lausanne, Switzerland
  • Entomological Institute, Czech Academy of Sciences, České Budějovice, Czech Republic
  • Národní museum, Prague, Czech Republic
  • Staatliches Museum für Naturkunde Lemberg, Ukraine
  • American Museum of Natural History, New York, USA
  • Albany Museum, Grahamstown, South Africa
  • Evolutionary Studies Institute, University of the Witwatersrand, Johannesburg, South Africa
  • Canterbury Museum, Christchurch, New Zealand

Field trips

  • Germany
  • New Zealand
  • Australia
  • Iran
  • South Africa
  • Vanuatu