Dr. Stephan Spiekman


Mitteltriassisische baumlebende Reptilienfauna


0711 / 8936-170
0711 / 8936-100


seit 2022

Postdoktorand, Naturkunde Museum Stuttgart, Arbeitsgruppe: Prof. Dr. Rainer Schoch


SNF Forschungstipendiat (Early Postdoc.Mobility Fellow), Natural History Museum London, Earth Sciences Department, Arbeitsgruppe: Dr. Susannah Maidment


3D Laborant (Wissenschaftlicher Mitarbeiter), Universität Zürich, Paläontologisches Institut und Museum


Doktorand, Universität Zürich, Paläontologisches Institut und Museum, Arbeitsgruppe: PD Dr. Torsten Scheyer


MSc. studium, Universität Leiden, Evolution, Biodiversity and Conservation


BSc. studium, Universität Leiden, Biology



  • Spiekman, S.N.F. and Klein, N., 2021. An enigmatic lower jaw from the Lower Muschelkalk (Anisian, Middle Triassic) of Winterswijk provides insights into dental configuration, tooth replacement and histology. Netherlands Journal of Geosciences 100: e17. https://doi.org/10.1017/njg.2021.12
  • Spiekman, S.N.F., Ezcurra, M.D., Butler, R.J., Fraser, N.C., Maidment, S.C.R., 2021. Pendraig milnerae, a new small-sized coelophysoid theropod from the Late Triassic of Wales. Royal Society Open Science 8(10): 210915. https://doi.org/10.1098/rsos.210915
  • Bakker, M.A.G., Van der Vos, W., De Jager, K., Chung, W-Y., Fowler, D.A., Dondorp, E., Spiekman, S.N.F., Chew, K-Y., Jiménez, R., Bickelmann, C., Kuratani, S., Blazek, R., Kondrashov, P., Renfree, M.B., Richardson, M.K., 2021. Selection on phalanx development in the evolution of the bird wing. Molecular Biology and Evolution 38(10): 4222-4237. https://doi.org/10.1093/molbev/msab150
  • Spiekman, S.N.F., Fraser, N.C., and Scheyer, T.M., 2021. A new phylogenetic hypothesis of Tanystropheidae (Diapsida, Archosauromorpha) and other “protorosaurs”, and its implications for the early evolution of stem archosaurs. PeerJ 9: e11143. https://doi.org/10.7717/peerj.11143


  • Spiekman, S.N.F., Neenan, J.M., Fraser, N.C., Fernandez, V., Rieppel, O., Nosotti, S., and Scheyer, T.M., 2020. The cranial morphology of Tanystropheus hydroides (Tanystropheidae, Archosauromorpha) as revealed by synchrotron microtomography. PeerJ 8: e10299. https://doi.org/10.7717/peerj.10299
  • Spiekman, S.N.F., Neenan, J.M., Fraser, N.C., Fernandez, V., Rieppel, O., Nosotti, S., and Scheyer, T.M., 2020. Aquatic habits and niche partitioning in the extraordinarily long-necked Triassic reptile Tanystropheus. Current Biology 30(19), 3889-3895. https://doi.org/10.1016/j.cub.2020.07.025
  • Miedema, F., Spiekman, S.N.F., Fernandez, V., Reumer, J.W.F., and Scheyer, T.M., 2020. Cranial morphology of the tanystropheid Macrocnemus bassanii unveiled using synchrotron microtomography. Scientific Reports 10: 12412. https://doi.org/10.1038/s41598-020-68912-4
  • Scheyer, T.M., Wang, W., Li, C., Miedema, F., and Spiekman, S.N.F., 2020. Osteological re-description of Macrocnemus fuyuanensis (Archosauromorpha, Tanystropheidae) from the Middle Triassic of China. Vertebrata Palasiatica 58(3): 169-187. https://doi.org/10.5167/uzh-191434
  • Scheyer, T.M., Spiekman, S.N.F., Sues, H.-D., Ezcurra, M.D., Butler, R.J., and Jones, M.E.H, 2020. Colobops, a juvenile rhynchocephalian reptile (Lepidosauromorpha), not a diminutive archosauromorph with an unusually strong bite. Royal Society Open Science 7(3). https://doi.org/10.1098/rsos.192179


  • Spiekman, S.N.F. and Scheyer, T. M., 2019. A taxonomic revision of the genus Tanystropheus (Archosauromorpha, Tanystropheidae). Palaeontologia Electronica 22.3.80, 1-46. https://doi.org/10.26879/1038  
  • Spiekman, S.N.F., Winkelhorst, H., Bleeker, R., Dorst, M., de Haan, R., and Voeten, D.F.A.E., 2019. Tanystropheids from the Winterswijkse Steengroeve – rare but recurring elements in the Vossenveld paleoassemblage. Grondboor & Hamer (Staringia 16), 73(5/6): 208-215. https://doi.org/10.5167/uzh-182944



  • Spiekman, S.N.F., 2018. A new specimen of Prolacerta broomi from the lower Fremouw Formation (Early Triassic) of Antarctica, its biogeographical implications and a taxonomic revision. Scientific Reports, 8(1):17996. https://doi.org/10.1038/s41598-018-36499-6
  • Werneburg, I. and Spiekman, S.N.F., 2018. Mammalian Embryology and Organogenesis. In: Zachos F, Asher R. (eds.). Handbook of Zoology: Mammalian Evolution, Diversity and Systematics. De Gruyter, Berlin. p. 59-116. https://doi.org/10.1515/9783110341553-004  


  • Spiekman, S.N.F. and Werneburg, I., 2017. Patterns in the bony skull development of marsupials: high variation in onset of ossification and conserved regions of bone contact. Scientific Reports 7: 43197. https://doi.org/10.1038/srep43197